His master's degree was on evolutionary genetics of salmonid fish.

Although, the development is not claimed to be of the same random nature as is by evolutionary genetics.

His main research field is the evolutionary genetics of mammals.

His areas of expertise are evolutionary genetics, host-parasite interactions and social evolution.

He is credited with founding the field of evolutionary genetics.

Innate social aptitudes of man: an approach from evolutionary genetics.

He has worked on a variety of subjects in evolutionary genetics.

Human evolutionary genetics and changes in the human genome over time.

He has made ample contributions to evolutionary genetics and population genetics.

He is considered a pioneer in the field of ecological and evolutionary genetics of plants.